What is the counter in Communications diagnostics?

All things groov RIO> IO> Diagnostic> Communication:

In an ongoing effort to define and debug the TCP Modbus issues we are having. Im poking all the things. Hey, we’re out here in IO/diagnostic, we have, 32 bit counters, neat right?

What are these counters telling me?
From a rough calculation, I would assert that in 49 days these counters are going to max out.
Then what? buffer overflow? System Crash? roll over to 0?

ARCNET (The LAN before time (its really REALLY OLD))
could not have had a 32 bit Ack counter in 1970. … so why are we tracking Arcnet Acks now?

I just want to know what this page is trying to tell me.